Shera, 21, one of the oldest African lions in the U.S., dies at Smithsonian’s National Zoo

Shera, an African lion housed at the Smithsonian’s National Zoo, has died at age 21, the zoo said. The animal, long familiar to visitors and staff, was one of the oldest lions in the United States.

The National Zoo confirmed Shera’s death on May 8 and noted that he sired eight cubs during his time at the institution. Those offspring have produced additional descendants now living at other accredited zoos across the country, giving Shera an extended legacy in the U.S. zoo community.

Shera’s presence at the National Zoo was part

Shera’s presence at the National Zoo was part of the institution’s larger animal-care and conservation work. The zoo has previously participated in managed breeding programs for lions and other species; Shera’s lineage reflects that network of accredited zoological institutions.

The Smithsonian’s National Zoo did not release detailed public information about the cause of Shera’s death in the initial announcement. Zoo staff typically review medical records and may share follow-up information about animal care and exhibit plans as appropriate.